“Diagnostic characteristics: Shiny shell with dense spiral striation above shell. The species differs from E. jucunda (which has similar sculpture) in terms of the shell shape and size (E. jucundior with five whorls and shell 19 mm in width vs. E. jucunda – with six and a half whorls and shell 17 mm in width).” (Liew et al., 2009)

“Remarks: The specimens in Laidlaw (1931) were collected by R. Hanitsch in 1899 in British North Borneo. According to the description by Laidlaw, these shells’ size and sculpture fit with the material we consider as E. jucundior. Some material in the RMNH and JJ collections from Balikpapan (Southeast Coast of Kalimantan) and Sarawak’ Fourth Division have similar spiral striation and shell shape but we do not include them in this Sabah Everettia revision. Our genetic data also show that E. jucundior is not closely related to other Sabah lowland Everettia species.” (Liew et al., 2009)

Author(s): Marzuki, Mohammad Effendi

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Liew et al. (2009) descriptions on Everettia jucundior – “Shell: medium-sized, rather thin, brownish, moderately elevated, outer whorls slightly shouldered or rounded. Periphery rounded. Above the periphery, shell shiny with densely placed spiral striation, c. 20–30 striae per mm. Below the periphery, shell has fine, densely placed spiral grooves. …number of whorls up to five and one-fifth…”

“Animal: black bands on either side of the tentacles, which are separated by a white band that occupies the space between the eye tentacles and extends downward to just below the eye tentacles. Grey smear on either side of the tail extending from the umbilicus region to the end of the tail. Other parts of the animal are clear from any markings. The mantle is black and covered by crowded tiny bright spots and followed by regularly spaced radial black bands in the last half whorl.” (Liew et al., 2009)

Author(s): Marzuki, Mohammad Effendi

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“Genitalia: maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to 11 mm. The penis, dart-sac, and vagina with thick muscular walls. Penis, vagina, genital opening, bursa copulatrix, and dart-sac arranged around the short genital atrium, with limited space between the opening of each of the genital structures. Bursa copulatrix short, about one-third of total dart-sac length.” (Liew et al., 2009)

Author(s): Marzuki, Mohammad Effendi

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Everettia jucundior – “Height up to 10.9 mm; width up to 19.5 mm; diameter of the first three whorls 0.9, 0.8, and 1.3 mm, respectively; height aperture up to 8.3 mm; width aperture up to 10.1 mm.” (Liew et al., 2009)

Author(s): Marzuki, Mohammad Effendi

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Type locality – “Sabah: Tawau District. Tawau Hills Park (4°24’N, 117°53’E)” leg. J. K. King/Dec. 2001 (Liew et al., 2009)

Other localities – “Sabah; Tawau, Batu Baturong, 100 m (alt.)” (Liew et al., 2009); “Sabah; Tawau, Tawau Hills Park” (Liew et al., 2009); “Sabah; Lahad Datu, Segama Valley, Kirk’s Cave” (Liew et al., 2009); “Sabah; Tabin Wildlife Reserve” (Liew et al., 2009); “Sabah; Danum Valley” (Liew et al., 2009); “Sabah; Kinabatangan, Batu Tomanggong Besar” (Liew et al., 2009); “Sabah; Kinabatangan, Batu Mawas” (Liew et al., 2009); “Sabah; Kinabatangan, Bod Tai” (Liew et al., 2009); “Sabah; Kinabatangan, Abai river” (Liew et al., 2009); “Sabah; Crocker Range, Keningau, Kampung Matou” (Liew et al., 2009); “Sabah; Semporna, Segarong Hills, Batu Tengar” (Liew et al., 2009)

“Distribution; Sabah: Kinabatangan, Tawau Hills Park, Danum valley.” (Liew et al., 2009)

Author(s): Marzuki, Mohammad Effendi

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“Habitat: Primary forest.” (Liew et al., 2009)

Author(s): Marzuki, Mohammad Effendi

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